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hembreeDan Hembree

Department of Earth and Planetary Sciences
703 Strong Hall
1621 Cumberland Ave
Knoxville, TN 37996-1410

Phone: 974-3788
Research website:

My research program involves the interaction of sedimentological and biological processes within the fields of paleopedology and ichnology. I combine these disciplines to study the evolution of modern and ancient soils, soil ecosystems, and landscapes as well as to demonstrate the broad environmental, climatic, and ecological applications of paleosols and ichnofossils, especially in evaluating the effect of long-term climate change on terrestrial ecosystems through Earth history. This involves not only the study of paleosols and continental trace fossils throughout geologic time, but also the experimental study of modern soils and the burrowing activities of extant terrestrial annelids, arthropods, amphibians, and reptiles.

My recent research projects have involved the study of the influence of climate change on ancient soils and soil ecosystems including those of the Pennsylvanian-Permian of southeast Ohio and West Virginia, as well as the Eocene to Pliocene of Colorado, Wyoming, southern Bolivia, and Ethiopia.

Selected Publications

  • Saylor, B.Z., Catena, A.M., Hembree, D.I., Anaya, F., Croft, D.A., 2022. Lithostratigraphy, chronostratigraphy, and sedimentary environments of the middle Miocene Quebrada Honda Basin in southern Bolivia and implications for Andean climate and uplift. Palaeogeography, Palaeoclimatology, Palaeoecology, v. 601, 111135.
  • Thacker, H.A., Hembree, D.I., 2021. Neoichnological study of burrowing darkling beetles (Coleoptera: Tenebrionidae) from larval to adult stages. Ichnos, v. 28(4), p. 290-308.
  • Hembree, D.I., McFadden, C.J., 2020. Analysis of climate and landscape change through the Pennsylvanian and Permian Monongahela and Dunkard Groups, Southeastern Ohio, USA. Journal of Sedimentary Environments, v. 5, p. 321-353.
  • Hembree, D.I., 2019. Burrows and ichnofabric produced by centipedes: modern and ancient examples. PALAIOS, v. 34, p. 468-489.
  • Kogler, S.J., Hembree, D.I., 2019. Influences of modern pedogenesis on paleoclimate estimates from Pennsylvanian and Permian paleosols, southeast Ohio. Journal of Sedimentary Research, v. 89, p. 227-241.
  • Hembree, D.I., Carnes, J.L., 2018. Response of soils and soil ecosystems to the Pennsylvanian–Permian climate transition in the upper fluvial plain of the Dunkard Basin, southeastern Ohio, USA. Geosciences, v. 8 (203), p. 1-35.
  • Hembree, D.I., Swaninger, E.S., 2018. Large Camborygma isp. in fluvial deposits of the Early Permian (Asselian) Dunkard Group, southeast Ohio, U.S.A. Palaeogeography, Palaeoclimatology, Palaeoecology, v. 491, p. 137-151.
  • Catena, A.M., Hembree, D.I., Saylor, B.Z., Croft, D.A., 2017. Paleosol and ichnofossil evidence for significant Neotropical habitat variation during the late middle Miocene (Serravallian). Palaeogeography, Palaeoclimatology, Palaeoecology, v. 487, p. 381-398.
  • Hembree, D.I., 2017. Neoichnology of tarantulas (Araneae: Theraphosidae): recognition of
    spider burrows in the fossil record. Palaeontologia Electronica 20.3.45A: 1-30.
  • Hembree, D.I., Smith, J.J., Buynevich, I.V., Platt, B.F., 2017. Neoichnology of semiarid environments: soils and burrowing animals of the Sonoran Desert, Arizona, U.S.A. PALAIOS, v. 32, p. 620-638.
  • Hembree, D.I., Bowen, J., 2017. Paleosols and ichnofossils of the Upper Pennsylvanian-Lower Permian Monongahela and Dunkard groups: a multi-proxy approach to unraveling complex variability in ancient terrestrial landscapes. PALAIOS, v. 32, p. 295-320.
  • Hembree, D.I., 2016. Using experimental neoichnology and quantitative analyses to improve the interpretation of continental trace fossils. Ichnos, v. 23, p. 262-297.
  • Hembree, D.I., Blair, M.G., 2016. A paleopedological and ichnological approach to interpreting spatial and temporal variability in Early Permian fluvial deposits of the Lower Dunkard Group, West Virginia, U.S.A. Palaeogeography, Palaeoclimatology, Palaeoecology, v. 454, p. 246-266.
  • Catena, A.M., Hembree, D.I., Saylor, B.Z., Anaya, F., Croft, D.A., 2016. Paleoenvironmental analysis of the Neotropical fossil mammal site of Cerdas, Bolivia (middle Miocene) based on ichnofossils and paleopedology. Palaeogeography, Palaeoclimatology, Palaeoecology, v.459, p. 423-439.
  • Suess, B., Wisshak, M., Mapes, R., Hembree, D., Landman, N., 2016. Microbial bioerosion of erratic sub-fossil Nautilus shells in a karstic cenote (Lifou, Loyalty Islands, New Caledonia). Ichnos, v. 23, 108-115.

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